We're getting a few things mixed up, so lets break this down into it's constituent parts.
The mechanical thrust or uplift of a pan is vertical. All the different movements and actions of the various parts, such as the upper and lower parts of the arm and the head, all resolve to remove any horizontal forces.
Put the train in motion and aerodynamic forces start to have an affect, but not really as much as some folk would imagine. No matter which way the pan is travelling, whether the knuckle is leading or trailing, the drag on the upper and lower parts of the arm are forcing one part up and the other part down. Just to illustrate this point, where the knuckle is leading the drag on the lower part of the arm will be attempting to force it up while the drag on the upper part will be attempting to force it down. Reverse the direction so that the knuckle is trailing and the forces are also reversed. However, the important point to consider here is that no matter which way round the pan is these forces acting on the pan as a consequence of drag will be the same.
In fact, in this particular area the BRBW high-speed pan performs better than the old Faiveley AMBR "bicycle frame" pan because the drag on each part of the arm is broadly balanced due to the broadly similar cross-section of each part. The old Faiveley pan almost certainly creates more drag with it's more complex upper section compared to the single arm used for the lower part.
So far so good. But where the BRBW pan caused problems at prototype stage was because of the design of the pan head with it's additional suspension unit. The aerodynamic drag on the carbon carriers and horns themselves was causing the pan head to lose contact with the wire with the knuckle trailing because the drag on the pan head itself was forcing it down and away from the contact wire. It was this problem that required the aerodynamic redesign of the head and the addition of aerofoils and tabs. As the speed increased and the aerodynamic drag on the head grew, so these modifications generated lift to counteract the effect and keep the pan head in contact with the wire.
As Old Timer suggests, the key is getting the amount of uplift right. Just as having too much uplift can cause bounce so can too little uplift, especially where the geometry of the OLE is perhaps a wee bit marginal. We have a couple of sections around Broxbourne where the pan loses contact because the contact wire changes height suddenly to accommodate a couple of bridges. Quite simply, when travelling at the full linespeed the pan cannot react to the height changes quickly enough and as a result loses contact.
Also, if I may be permitted to clear up a couple of other associated points.
The proposed APT-S production train required two motor coaches in order to achieve the high speeds required. However, these were required to be in the centre of the train primarily because of the reluctance to have a 25kV "bus" running the length of the train. The use of two pans was tested at speeds exceeding 125mph using a temporary HST formation, as described in the webpage I linked to earlier, and the performance was found to be acceptable. In addition, the original testing of the BRBW pan was done by hooking Lab 6 "Prometheus" onto the back on an electric train and running it up and down the WCML between London and Carlisle. Even then "...the pantograph's performance at the rear of the train as the second pantograph was remarkable. It was as good as the Faiveley when leading and our pan was running on an already moving contact wire."
I would usually defer to Old Timer's greater experience and depth of knowledge, but I'm not sure that the theory that a snagged pan would be likely to incur less damage travelling one way compared to the other holds up in practice. Clearly operational experience must have demonstrated that any advantage was small as the practice of fitting two pans to locos was short-lived and not applied to EMU's.
Now it may just be the customary practice of Virgin to run a particular way around or the pans on the Cl390's may have been specifically tuned to alter the amount of uplift, but that is nothing to do with any shortcomings of the pan design itself. One or two minor operating advantages have been pointed out to me by another member by PM, but those alone still don't really convince me that there is any significant advantage to running the pan in one direction or the other.
Nor, for that matter, why it should be generating quite so much interest or discussion.
O L Leigh