I believe there are very few people in the country who will know where the revenue for a Nantwich to Wrexham ticket goes. The formula for calculating ORCATS payments is a closely guarded secret. We know very little about it.
I can guess that it is highly likely that the majority of revenue goes to ATW, as a quick look at the timetable shows the majority of the fastest journey opportunities involving 3 ATW trains, but there are some of the fastest journey opportunities that involve either VT or WS for a significant portion, so they'll get a fair proportion too. Other TOCs are unlikely to get much. All applicable TOCs are required to convey customers holding such tickets on valid routes as permitted in the Routeing Guide, subject to ticket conditions & restrictions regardless of revenue allocation.
If any TOC feels that they are not getting a fair proportion of any particular flow, they can request for an audit to be carried out. However this may be costly and it is unknown (that I know of) who pays for the cost of such an audit.
See other threads - this has cropped up several times recently, try searching the forum for ORCATS.
I'd really like to know, for example, how much ORCATS revenue the 2033 King's Cross - Nottingham (change @ Grantham) gets versus the 2030 St Pancras - Nottingham (direct but arrives Nottingham 2 mins later and overall 5 mins slower). I suspect most people here would go for the faster option as it's a vastly superior train and service with better on-board facilities, catering etc, and most here would know it's a simple case of walking from one side of a platform to another, and the chance to get fresh air for a few mins during the journey is usually welcome. But you can guarantee any elderly, disabled, or families using the train would want to get the direct train and wouldn't realise that they're getting a rattling mid-range DMU for such a journey (and they might not even care!). The irony is that those coming from the London Underground station (ie, the majority of passengers I reckon) would do a lot less walking at King's Cross and at Grantham combined than the St Pancras people would! But would ORCATS know any of this? I'd hope that ORCATS would give more money to the faster journey, but I suspect probably not as the direct train is only 5 minutes slower and it is weighted toward direct trains. But we can only guess!