Having got hold of a copy of MacDermot I find there is a letter by Gooch on the subject, complete with cross-sectional diagram of the boiler, which explains in some detail these engines' tendency to prime; basically there was very little steam space above the surface of the water in the boiler barrel and what there was was mostly filled by the main steam pipe, so when the engine was using a lot of steam and the water was boiling hard, the rush of steam through this restricted space would pick up water and blow it along with it to the steam collector. He then goes on to describe modifications that would increase the steam space, and hopes that when this is done "we will be able to run with a safe level of water in the boiler".
He says the reason this is difficult at present is that if the driver is running fast and then shuts off steam for a station, the water level drops and exposes the firebox crown, so instead the driver has to reverse the motion while keeping steam on. Unfortunately that's all he does say, and it still leaves the puzzle of why shutting off steam should cause the water level to drop.
My best guess so far is that when the engine was running fast, continually using a lot of steam and then replenishing the water as required using the axle-driven feed pump, it was possible to run the quantity of water in the boiler down to below the normal static minimum level while still maintaining an adequate "dynamic" level; the boiler could end up in an equilibrium condition where the rush of steam towards the firebox end was carrying enough rush of water with it to maintain a level above the firebox crown and an indication in the gauge glass before it ran back again into the main part of the boiler, even though the static level of the same quantity of boiler water would be below the firebox crown (but higher in the main part of the boiler). Stopping the flow of steam would stop the rush of water and allow it to settle at an inadequate static level. So they would have to reduce the steam flow gradually while turning on the feed pump in order to return to a statically safe condition.